Although the Wye certainly suited the fish that were imported in the 1890's, I think the circumstances that brought about their introduction and subsequent release into the river proper from Ashford Dam, held paramount importance.
Let me explain:
The original Wye fish came from a watershed and hatchery very different from later imports. Most significant was that they were spring spawning, where as the Shasta's from the McCloud River, spawn well before Christmas. This makes them ultimately successful because their eggs do not sit in the gravel all winter, with their biological clocks turned off due to low water temperatures. Our spring spawning fish hatch in no time and although they appear from very small eggs, they swim-up into water that has developed, as far as zoo plankton is concerned.
I also believe that the broodstock played a vital role too. Surely, as fish farming was in its infancy, brood fish were caught up rather than being held. This gave their babies a good chance of inheriting the genes that would help them recognise spawning cues, when the time came.
Four years into their life in the lake at Ashford they were able to get out following periods of exceptionally high water. Coming from water with a temperature and pH very much like the Wye (Frost) and being liberated when they were all sexually mature (and understanding spawning cues) they spawned and the rest is history.
I wasn't ever able to ascertain whether the Misbourne and Chess rainbows were spring spawning through my correspondence with F.J.Taylor but I do know that the river and a lake on the Misbourne (name escapes me) were stocked with Shasta's from the local fish farm after a severe drought. Those fish never spawned and died out.
It is possible for Shasta babies, released accidentally from fish farms, to spawm when they get older, as is the case on the nearby River Dove. Historically these fish die out after a few generations but I don't know why.
Occasionally we see fish that have such tremendous scale adhesion, they are almost like enamel.
But the average fish are much most spotty. Here's Gareth with a cock fish...